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I don't believe you can change a races base gravity tolerance through research at all.
After researching genome sequence, I always found all the others unlocked, so I thought that is the only prerequisite. By the way, you can have a look to the tech tree posted by Turmoil.
I'm still with 5.60, so I'm afraid I can't help you, sorry. Just can tell you that with 5.60 it's available as soon as you have genome sequence. Maybe Steve removed it in 6.X?